| Citation | Distinguished for his comparative studies of the behavioural ecology of large mammals. His early work demonstrated that the social behaviour of primates is markedly influenced by the temporal and spatial distribution of their food. Subsequently he has vigorously led a team in a remarkable 15 year study of the red deer on the island of Rhum. With skill and rigour he has shown how the differences between the sexes in mortality and feeding behaviour can be related to the highly polygamous mating system. He has also provided strong evidence that the amount of paternal care invested in offspring depends on the sex of the offspring as well as the condition of the mother. This major study of the red deer is widely regarded as a classic. By the depth of his own empirical work and the breadth of his comparative surveys, he has greatly increased our understanding of how sexual and parental behaviour have evolved and may be related to the ecological conditions in which animals live. |