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  <PersonName label="Authorised form of name" urlencoded="Pennycuick%3b+Colin+James+(1933+-+2019)" urlpathencoded="Pennycuick;%20Colin%20James%20(1933%20-%202019)">Pennycuick; Colin James (1933 - 2019)</PersonName>
  <Surname label="Surname" urlencoded="Pennycuick" urlpathencoded="Pennycuick">Pennycuick</Surname>
  <Forenames label="Forenames" urlencoded="Colin+James" urlpathencoded="Colin%20James">Colin James</Forenames>
  <PreTitle label="PreTitle" urlencoded="" urlpathencoded=""></PreTitle>
  <Title label="Title" urlencoded="" urlpathencoded=""></Title>
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  <Dates label="Dates" urlencoded="1933+-+2019" urlpathencoded="1933%20-%202019">1933 - 2019</Dates>
  <Nationality label="Nationality" urlencoded="British" urlpathencoded="British">British</Nationality>
  <Gender label="Gender" urlencoded="Male" urlpathencoded="Male">Male</Gender>
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  <PlaceOfBirth label="Place of birth" urlencoded="Virginia+Water%2c+Surrey%2c+England%2c+United+Kingdom" urlpathencoded="Virginia%20Water,%20Surrey,%20England,%20United%20Kingdom">Virginia Water, Surrey, England, United Kingdom </PlaceOfBirth>
  <DateOfBirth label="Date of birth" urlencoded="11%2f06%2f1933" urlpathencoded="11/06/1933">11/06/1933</DateOfBirth>
  <Born label="Born" urlencoded="19330611" urlpathencoded="19330611">19330611</Born>
  <PlaceOfDeath label="Place of death" urlencoded="" urlpathencoded=""></PlaceOfDeath>
  <DateOfDeath label="Date of death" urlencoded="09%2f12%2f2019" urlpathencoded="09/12/2019">09/12/2019</DateOfDeath>
  <Died label="Died" urlencoded="20191209" urlpathencoded="20191209">20191209</Died>
  <DatesAndPlaces label="DatesAndPlaces" urlencoded="" urlpathencoded=""></DatesAndPlaces>
  <Address label="Address" urlencoded="" urlpathencoded=""></Address>
  <Occupation label="Occupation" urlencoded="Zoologist" urlpathencoded="Zoologist">Zoologist</Occupation>
  <ResearchField label="Research field" urlencoded="Biology" urlpathencoded="Biology">Biology  </ResearchField>
  <ResearchField label="Research field" urlencoded="Zoology" urlpathencoded="Zoology">Zoology  </ResearchField>
  <ResearchField label="Research field" urlencoded="Ornithology" urlpathencoded="Ornithology">Ornithology  </ResearchField>
  <ResearchField label="Research field" urlencoded="Flight" urlpathencoded="Flight">Flight  </ResearchField>
  <Activity label="Activity" urlencoded="Education%3a%0aLanesborough+Prep+School%2c+Guildford+(1943-1947)%3b+Wellington+College%2c+Berkshire+(1947-1951)%3b+Merton+College%2c+University+of+Oxford%2c+BSc+in+Natural+Sciences+-+Zoology+(1951-1955)%3b+Peterhouse%2c+University+of+Cambridge%2c+PhD+in+animal+behaviour+(1962)%0aCareer%3a%0aNational+Service+in+Royal+Airforce+(1955-1957)%3b+Qualified+as+RAF+pilot+(1957)%3b+Research+Assistant+to+Dr+Thorpe%2c+Department+of+Animal+Behavious%2c+Madingley+(1960-1964)%3b+Assistant+Lecturer%2fLecturer++in+Zoology%2c+University+of+Bristol+(1964-1968)%3b+Lecturer+in+Zoology%2c+University+of+Nairobi%2c+Kenya+(on+secondment+1968-1971)%3b+Deputy+Directpr%2c+Serengeti+Research+Institute%2c+Tanzania+(1971-1973)%3b+Lecturer+in+Zoology+(1973-1975)+Reader+in+Zoology+(1975-1983)+University+of+Bristol%3b+Maytag+Professor+of+Ornithology%2c+University+of+Miami%2c+United+States+of+America+(1983-1992)%3b+Animal+Ecology+Group%2c+Lund+University+(1992-)%3b++Wildlife+and+Wetlands+Trust%2c+Gloucestershire%3b+research+professor+in+zoology+(1993-1997)%3b+senior+research+fellow+(1997-)%2c+University+of+Bristol%0aMemberships%3a%0aCorresponding+Fellow+of+American+Ornithologists%27+Union+(1975)%3b+Honorary+Companion+of+the+Royal+Aeronautical+Society+(1994)%0aMedals%2fAwards%3a%0aHonorary+doctorate+from+Lund+University+(1996)" urlpathencoded="Education:%0aLanesborough%20Prep%20School,%20Guildford%20(1943-1947);%20Wellington%20College,%20Berkshire%20(1947-1951);%20Merton%20College,%20University%20of%20Oxford,%20BSc%20in%20Natural%20Sciences%20-%20Zoology%20(1951-1955);%20Peterhouse,%20University%20of%20Cambridge,%20PhD%20in%20animal%20behaviour%20(1962)%0aCareer:%0aNational%20Service%20in%20Royal%20Airforce%20(1955-1957);%20Qualified%20as%20RAF%20pilot%20(1957);%20Research%20Assistant%20to%20Dr%20Thorpe,%20Department%20of%20Animal%20Behavious,%20Madingley%20(1960-1964);%20Assistant%20Lecturer/Lecturer%20%20in%20Zoology,%20University%20of%20Bristol%20(1964-1968);%20Lecturer%20in%20Zoology,%20University%20of%20Nairobi,%20Kenya%20(on%20secondment%201968-1971);%20Deputy%20Directpr,%20Serengeti%20Research%20Institute,%20Tanzania%20(1971-1973);%20Lecturer%20in%20Zoology%20(1973-1975)%20Reader%20in%20Zoology%20(1975-1983)%20University%20of%20Bristol;%20Maytag%20Professor%20of%20Ornithology,%20University%20of%20Miami,%20United%20States%20of%20America%20(1983-1992);%20Animal%20Ecology%20Group,%20Lund%20University%20(1992-);%20%20Wildlife%20and%20Wetlands%20Trust,%20Gloucestershire;%20research%20professor%20in%20zoology%20(1993-1997);%20senior%20research%20fellow%20(1997-),%20University%20of%20Bristol%0aMemberships:%0aCorresponding%20Fellow%20of%20American%20Ornithologists'%20Union%20(1975);%20Honorary%20Companion%20of%20the%20Royal%20Aeronautical%20Society%20(1994)%0aMedals/Awards:%0aHonorary%20doctorate%20from%20Lund%20University%20(1996)">Education:
Lanesborough Prep School, Guildford (1943-1947); Wellington College, Berkshire (1947-1951); Merton College, University of Oxford, BSc in Natural Sciences - Zoology (1951-1955); Peterhouse, University of Cambridge, PhD in animal behaviour (1962)
Career:
National Service in Royal Airforce (1955-1957); Qualified as RAF pilot (1957); Research Assistant to Dr Thorpe, Department of Animal Behavious, Madingley (1960-1964); Assistant Lecturer/Lecturer  in Zoology, University of Bristol (1964-1968); Lecturer in Zoology, University of Nairobi, Kenya (on secondment 1968-1971); Deputy Directpr, Serengeti Research Institute, Tanzania (1971-1973); Lecturer in Zoology (1973-1975) Reader in Zoology (1975-1983) University of Bristol; Maytag Professor of Ornithology, University of Miami, United States of America (1983-1992); Animal Ecology Group, Lund University (1992-);  Wildlife and Wetlands Trust, Gloucestershire; research professor in zoology (1993-1997); senior research fellow (1997-), University of Bristol
Memberships:
Corresponding Fellow of American Ornithologists' Union (1975); Honorary Companion of the Royal Aeronautical Society (1994)
Medals/Awards:
Honorary doctorate from Lund University (1996)</Activity>
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  <OtherInfo label="OtherInfo" urlencoded="Colin+Pennycuick+was+a+leading+authority+on+the+flight+of+birds+and+bats%2c+and+of+bird+navigation+and+migration.+Amongst+his+many+contributions%2c+were+advancing+our+understanding+of+the+relationship+between+the+power+of+flight+muscles%2c+wingbeat+frequency+and+the+power+density+of+mitochondria+%e2%80%94+the+%e2%80%98powerhouses%e2%80%99+of+cells.+Colin+also+shed+light+on+species-specific+phenomena+such+as+gust+soaring+in+albatrosses.%0a%0aHe+was+known+for+his+innovative+data+collection+from+controlled+experiments+%e2%80%94+he+pioneered+the+use+of+wind+tunnels+for+flight+studies%2c+for+example+%e2%80%94+and+observations%2c+including+those+made+using+a+powered+glider.+He+also+developed+the+ornithodolite%2c+a+portable+instrument+for+real-time+computer+recording+of+azimuth%2c+elevation+and+range+of+birds+in+flight.%0a%0aColin%e2%80%99s+%e2%80%98Flight%e2%80%99+software+models+the+mechanics+of+bird+flight%2c+covering+flapping+and+gliding+flight+and+long-distance+migration.+His+textbook%2c+Modelling+the+Flying+Bird+(2008)%2c+complemented+his+software%2c+setting+out+the+principles+of+aeronautical+engineering+and+how+they+may+be+adapted+to+exploring+biological+problems+of+bird+flight.%0a%0aProfessor+Colin+Pennycuick+FRS+died+on+9+December+2019." urlpathencoded="Colin%20Pennycuick%20was%20a%20leading%20authority%20on%20the%20flight%20of%20birds%20and%20bats,%20and%20of%20bird%20navigation%20and%20migration.%20Amongst%20his%20many%20contributions,%20were%20advancing%20our%20understanding%20of%20the%20relationship%20between%20the%20power%20of%20flight%20muscles,%20wingbeat%20frequency%20and%20the%20power%20density%20of%20mitochondria%20%e2%80%94%20the%20%e2%80%98powerhouses%e2%80%99%20of%20cells.%20Colin%20also%20shed%20light%20on%20species-specific%20phenomena%20such%20as%20gust%20soaring%20in%20albatrosses.%0a%0aHe%20was%20known%20for%20his%20innovative%20data%20collection%20from%20controlled%20experiments%20%e2%80%94%20he%20pioneered%20the%20use%20of%20wind%20tunnels%20for%20flight%20studies,%20for%20example%20%e2%80%94%20and%20observations,%20including%20those%20made%20using%20a%20powered%20glider.%20He%20also%20developed%20the%20ornithodolite,%20a%20portable%20instrument%20for%20real-time%20computer%20recording%20of%20azimuth,%20elevation%20and%20range%20of%20birds%20in%20flight.%0a%0aColin%e2%80%99s%20%e2%80%98Flight%e2%80%99%20software%20models%20the%20mechanics%20of%20bird%20flight,%20covering%20flapping%20and%20gliding%20flight%20and%20long-distance%20migration.%20His%20textbook,%20Modelling%20the%20Flying%20Bird%20(2008),%20complemented%20his%20software,%20setting%20out%20the%20principles%20of%20aeronautical%20engineering%20and%20how%20they%20may%20be%20adapted%20to%20exploring%20biological%20problems%20of%20bird%20flight.%0a%0aProfessor%20Colin%20Pennycuick%20FRS%20died%20on%209%20December%202019.">Colin Pennycuick was a leading authority on the flight of birds and bats, and of bird navigation and migration. Amongst his many contributions, were advancing our understanding of the relationship between the power of flight muscles, wingbeat frequency and the power density of mitochondria — the ‘powerhouses’ of cells. Colin also shed light on species-specific phenomena such as gust soaring in albatrosses.

He was known for his innovative data collection from controlled experiments — he pioneered the use of wind tunnels for flight studies, for example — and observations, including those made using a powered glider. He also developed the ornithodolite, a portable instrument for real-time computer recording of azimuth, elevation and range of birds in flight.

Colin’s ‘Flight’ software models the mechanics of bird flight, covering flapping and gliding flight and long-distance migration. His textbook, Modelling the Flying Bird (2008), complemented his software, setting out the principles of aeronautical engineering and how they may be adapted to exploring biological problems of bird flight.

Professor Colin Pennycuick FRS died on 9 December 2019.</OtherInfo>
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  <DateOfElection label="Date of election" urlencoded="15%2f03%2f1990" urlpathencoded="15/03/1990">15/03/1990</DateOfElection>
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  <Source label="Source" urlencoded="Sources%3a%0ahttps%3a%2f%2fwww.theguardian.com%2fscience%2f2020%2ffeb%2f24%2fcolin-pennycuick-obituary+%0ahttps%3a%2f%2froyalsociety.org%2fpeople%2fcolin-pennycuick-12073%2f" urlpathencoded="Sources:%0ahttps://www.theguardian.com/science/2020/feb/24/colin-pennycuick-obituary%20%0ahttps://royalsociety.org/people/colin-pennycuick-12073/">Sources:
https://www.theguardian.com/science/2020/feb/24/colin-pennycuick-obituary 
https://royalsociety.org/people/colin-pennycuick-12073/</Source>
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National Council on Archives, Rules for the Construction of Personal, Place and Corporate Names, 1997</Conventions>
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Pmab</Creator>
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