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  <PersonName label="Authorised form of name" urlencoded="Barlow%3b+Horace+Basil+(1921+-+2020)" urlpathencoded="Barlow;%20Horace%20Basil%20(1921%20-%202020)">Barlow; Horace Basil (1921 - 2020)</PersonName>
  <Surname label="Surname" urlencoded="Barlow" urlpathencoded="Barlow">Barlow</Surname>
  <Forenames label="Forenames" urlencoded="Horace+Basil" urlpathencoded="Horace%20Basil">Horace Basil</Forenames>
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  <Dates label="Dates" urlencoded="1921+-+2020" urlpathencoded="1921%20-%202020">1921 - 2020</Dates>
  <Nationality label="Nationality" urlencoded="British" urlpathencoded="British">British</Nationality>
  <Gender label="Gender" urlencoded="Male" urlpathencoded="Male">Male</Gender>
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  <PlaceOfBirth label="Place of birth" urlencoded="England%2c+United+Kingdom" urlpathencoded="England,%20United%20Kingdom">England, United Kingdom </PlaceOfBirth>
  <DateOfBirth label="Date of birth" urlencoded="08%2f12%2f1921" urlpathencoded="08/12/1921">08/12/1921</DateOfBirth>
  <Born label="Born" urlencoded="19211208" urlpathencoded="19211208">19211208</Born>
  <PlaceOfDeath label="Place of death" urlencoded="" urlpathencoded=""></PlaceOfDeath>
  <DateOfDeath label="Date of death" urlencoded="05%2f07%2f2020" urlpathencoded="05/07/2020">05/07/2020</DateOfDeath>
  <Died label="Died" urlencoded="20200705" urlpathencoded="20200705">20200705</Died>
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  <Address label="Address" urlencoded="" urlpathencoded=""></Address>
  <Occupation label="Occupation" urlencoded="Neuroscientist" urlpathencoded="Neuroscientist">Neuroscientist</Occupation>
  <ResearchField label="Research field" urlencoded="Neuroscience" urlpathencoded="Neuroscience">Neuroscience</ResearchField>
  <ResearchField label="Research field" urlencoded="Vision" urlpathencoded="Vision">Vision</ResearchField>
  <ResearchField label="Research field" urlencoded="Physiological+optics" urlpathencoded="Physiological%20optics">Physiological optics</ResearchField>
  <Activity label="Activity" urlencoded="Education%3a+%0aWinchester+College%3b+Harvard+University%2c+MD+(1946)+%0aCareer%3a+%0aFellow+of+Trinity+College%2c+University+of+Cambridge%3b+Royal+Society+Research+Professor+of+Physiology+at+the+University+of+Cambridge%3b+University+of+California+at+Berkeley+(1964-1973)%3b+University+of+Cambridge+(1973-2020)+%0aMedals%2fAwards%3a+%0aAustralia+Prize+1993%2c+Swartz+Prize+for+Theoretical+and+Computational+Neuroscience+2009%3b+Ken+Nakayama+Prize+2016" urlpathencoded="Education:%20%0aWinchester%20College;%20Harvard%20University,%20MD%20(1946)%20%0aCareer:%20%0aFellow%20of%20Trinity%20College,%20University%20of%20Cambridge;%20Royal%20Society%20Research%20Professor%20of%20Physiology%20at%20the%20University%20of%20Cambridge;%20University%20of%20California%20at%20Berkeley%20(1964-1973);%20University%20of%20Cambridge%20(1973-2020)%20%0aMedals/Awards:%20%0aAustralia%20Prize%201993,%20Swartz%20Prize%20for%20Theoretical%20and%20Computational%20Neuroscience%202009;%20Ken%20Nakayama%20Prize%202016">Education: 
Winchester College; Harvard University, MD (1946) 
Career: 
Fellow of Trinity College, University of Cambridge; Royal Society Research Professor of Physiology at the University of Cambridge; University of California at Berkeley (1964-1973); University of Cambridge (1973-2020) 
Medals/Awards: 
Australia Prize 1993, Swartz Prize for Theoretical and Computational Neuroscience 2009; Ken Nakayama Prize 2016
</Activity>
  <Relationships label="Relationships" urlencoded="Son+of+the+civil+servant+Sir+Alan+Barlow+and+his+wife+Lady+Nora+(n%c3%a9e+Darwin)%2c+great+grandson+of+Charles+Darwin+FRS%3b+married+1.+(1954)+Ruthala+Salaman%2c+2.+(1980)+Miranda+Weston-Smith%3b+professional+collaboration+with+Richard+Fitzhugh%2c+Stephen+Kuffler%2c+William+Levick" urlpathencoded="Son%20of%20the%20civil%20servant%20Sir%20Alan%20Barlow%20and%20his%20wife%20Lady%20Nora%20(n%c3%a9e%20Darwin),%20great%20grandson%20of%20Charles%20Darwin%20FRS;%20married%201.%20(1954)%20Ruthala%20Salaman,%202.%20(1980)%20Miranda%20Weston-Smith;%20professional%20collaboration%20with%20Richard%20Fitzhugh,%20Stephen%20Kuffler,%20William%20Levick">Son of the civil servant Sir Alan Barlow and his wife Lady Nora (née Darwin), great grandson of Charles Darwin FRS; married 1. (1954) Ruthala Salaman, 2. (1980) Miranda Weston-Smith; professional collaboration with Richard Fitzhugh, Stephen Kuffler, William Levick</Relationships>
  <OtherInfo label="OtherInfo" urlencoded="Horace+Barlow+was+best+known+for+his+analysis+of+the+way+that+visual+information+is+encoded+as+significant+nerve+signals.+He+was+the+first+to+show+(in+amphibians)+the+fundamental+organisation+of+optic+nerve+fibres%2c+namely+that+each+is+connected+to+a+clump+of+receptors+that+excite%2c+surrounded+by+a+ring+that+inhibits.+With+Richard+Fitzhugh+and+Stephen+Kuffler%2c+he+analysed+how+this+organization+changes+in+dark+adaptation.%0a%0aHorace%e2%80%99s+work+on+human+vision+greatly+strengthened+the+modern+concept+of+threshold+as+signal+detection+in+noisy+channels%2c+and+he+measured+the+quantum+efficiency+of+seeing+in+various+conditions.+With+William+Levick%2c+he+showed+that+rabbits+have+optic+nerves+that+signal+not+position+but+direction+of+movement%2c+and+by+ingenious+experiments+analysed+the+neural+mechanism+by+which+specific+movement+is+encoded.%0a%0aDr+Horace+Barlow+FRS+died+on+5+July+2020." urlpathencoded="Horace%20Barlow%20was%20best%20known%20for%20his%20analysis%20of%20the%20way%20that%20visual%20information%20is%20encoded%20as%20significant%20nerve%20signals.%20He%20was%20the%20first%20to%20show%20(in%20amphibians)%20the%20fundamental%20organisation%20of%20optic%20nerve%20fibres,%20namely%20that%20each%20is%20connected%20to%20a%20clump%20of%20receptors%20that%20excite,%20surrounded%20by%20a%20ring%20that%20inhibits.%20With%20Richard%20Fitzhugh%20and%20Stephen%20Kuffler,%20he%20analysed%20how%20this%20organization%20changes%20in%20dark%20adaptation.%0a%0aHorace%e2%80%99s%20work%20on%20human%20vision%20greatly%20strengthened%20the%20modern%20concept%20of%20threshold%20as%20signal%20detection%20in%20noisy%20channels,%20and%20he%20measured%20the%20quantum%20efficiency%20of%20seeing%20in%20various%20conditions.%20With%20William%20Levick,%20he%20showed%20that%20rabbits%20have%20optic%20nerves%20that%20signal%20not%20position%20but%20direction%20of%20movement,%20and%20by%20ingenious%20experiments%20analysed%20the%20neural%20mechanism%20by%20which%20specific%20movement%20is%20encoded.%0a%0aDr%20Horace%20Barlow%20FRS%20died%20on%205%20July%202020.">Horace Barlow was best known for his analysis of the way that visual information is encoded as significant nerve signals. He was the first to show (in amphibians) the fundamental organisation of optic nerve fibres, namely that each is connected to a clump of receptors that excite, surrounded by a ring that inhibits. With Richard Fitzhugh and Stephen Kuffler, he analysed how this organization changes in dark adaptation.

Horace’s work on human vision greatly strengthened the modern concept of threshold as signal detection in noisy channels, and he measured the quantum efficiency of seeing in various conditions. With William Levick, he showed that rabbits have optic nerves that signal not position but direction of movement, and by ingenious experiments analysed the neural mechanism by which specific movement is encoded.

Dr Horace Barlow FRS died on 5 July 2020.</OtherInfo>
  <ROYAL_SOCIETY_ACTIVITY label="ROYAL_SOCIETY_ACTIVITY" urlencoded="" urlpathencoded=""></ROYAL_SOCIETY_ACTIVITY>
  <RSActivity label="RSActivity" urlencoded="Royal+Society+roles%3a%0aCouncil%3a+1989-1990%0aMedals+and+prizes%3a%0aRoyal+Medal+1993+%0aLectures%3a%0aFerrier+1980" urlpathencoded="Royal%20Society%20roles:%0aCouncil:%201989-1990%0aMedals%20and%20prizes:%0aRoyal%20Medal%201993%20%0aLectures:%0aFerrier%201980">Royal Society roles:
Council: 1989-1990
Medals and prizes:
Royal Medal 1993 
Lectures:
Ferrier 1980 </RSActivity>
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  <DateOfElection label="Date of election" urlencoded="20%2f03%2f1969" urlpathencoded="20/03/1969">20/03/1969</DateOfElection>
  <AgeAtElection label="Age at election" urlencoded="47" urlpathencoded="47">47</AgeAtElection>
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  <Source label="Source" urlencoded="Sources%3a%0ahttps%3a%2f%2froyalsociety.org%2fpeople%2fhorace-barlow-11035%2f%3b+Wikipedia%0aReferences%3a%0aDavid+Burr%2c+%27Interview+with+Horace+Barlow%27+in+Current+Biology+volume+18%2c+issue+12%2c+June+2008+https%3a%2f%2fdoi.org%2f10.1016%2fj.cub.2008.04.005+(accessed+6%2f7%2f2020)" urlpathencoded="Sources:%0ahttps://royalsociety.org/people/horace-barlow-11035/;%20Wikipedia%0aReferences:%0aDavid%20Burr,%20'Interview%20with%20Horace%20Barlow'%20in%20Current%20Biology%20volume%2018,%20issue%2012,%20June%202008%20https://doi.org/10.1016/j.cub.2008.04.005%20(accessed%206/7/2020)">Sources:
https://royalsociety.org/people/horace-barlow-11035/; Wikipedia
References:
David Burr, 'Interview with Horace Barlow' in Current Biology volume 18, issue 12, June 2008 https://doi.org/10.1016/j.cub.2008.04.005 (accessed 6/7/2020)</Source>
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National Council on Archives, Rules for the Construction of Personal, Place and Corporate Names, 1997</Conventions>
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