| Citation | Distinguished by his contributions to various aspects of plant cell biology, where he has developed a highly productive approach involving fine-structural, physiological and biochemical techniques. Discovered the stromacentre in Avena plastids, and analysed the geometry of the prolamellar body of etiolated plastids and of its conversion into the thylakoid system during greening. With Pate has studied the structure, development, function and taxonomic distribution of transfer cells, making here an important new contribution to physiological plant anatomy, since the significance of these cells in providing a short-range, high intensity translocation system for solutes has hitherto escaped recognition. Latterly has begun a study of the architecture of certain algal cells using stereology as a means of three-dimensional reconstruction, and has shown that the unique plant wall polymer, sporpollenin, hitherto known only from the spores of vascular plants, is produced by certain green algae. |