Citation | Emeritus Professor of Reproductive Sciences, Department of Physiology, Development and Neuroscience, University of Cambridge
Johnson's seminal scientific contribution was the discovery and analysis of cellular polarization during early mammalian development. He showed that this event initiated the first lineage segregation: one lineage formed the outer implanting layer of the placenta while the fetal body developed form the other. Recent techniques have permitted further understanding of this vital and decisive moment, and they all depend and build on his foundations. He also contributed to human reproductive sciences with his work leading to change in clinical practice.
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