Citation | Professor Ewens has made major contributions in three main areas of population genetics theory and its applications. His formula describing the properties of a sample of genes from a selectively neutral locus have been highly influential in evolutionary genetics, probability theory and combinatorial theory. He established and proved the modern version of Fisher's fundamental theorem of natural selection and the evolutionary opitimality principle flowing from it. More recently, in human genetics he developed the transmission disequilibrium test which has become widely used in searching for genes involved with complex diseases |